Overview

The most well-known archaic human, with hundreds of fossils and a fully sequenced genome. Characterized by a robust, cold-adapted body, large brain (often exceeding modern humans), prominent brow ridges, and a projecting midface. Produced sophisticated Mousterian stone tools, used fire, buried their dead (Shanidar Cave, La Ferrassie), cared for injured individuals, and may have created symbolic art. Interbred with H. sapiens — 1-4% of modern non-African DNA is Neanderthal. Multiple lines of evidence suggest complex cognitive and social abilities.

Key Fossils

Neanderthal 1, La Chapelle-aux-Saints, Shanidar 1-9, La Ferrassie, Kebara 2, Altai Neanderthal

Brain Anatomy

Endocast-Based Estimates

Brain region sizes are estimated from endocasts (internal skull casts), not direct brain observation. These are approximations with inherent uncertainty.
Frontal
508 cc
Parietal
348 cc
Temporal
290 cc
Occipital
189 cc
Cerebellar
116 cc
RegionFunctional Implicationsvs. Modern Human
Frontal Large frontal lobes but relatively less prefrontal than sapiens; strong executive function Similar total volume but different internal organization; less globular
Parietal Large parietal lobes; excellent visuospatial skills for Mousterian tool-making Less expanded parietal than sapiens; different shape
Temporal Well-developed temporal lobes; social cognition, possible language Similar volume; less temporal pole expansion
Occipital Larger proportional occipital than sapiens; more visual cortex allocation Larger proportionally; more visual processing emphasis
Cerebellar Moderate cerebellar development Slightly smaller proportionally

Tools & Technology

Associated Tool Traditions

Mousterian

300 KYA – 30 KYA

Middle Paleolithic prepared-core technology primarily associated with Neanderthals. Levallois technique produces predetermined flake shapes. Includes points, scrapers, and notched tools.

Chatelperronian

44 KYA – 36 KYA

A transitional Upper Paleolithic tradition found in France and Spain, attributed to late Neanderthals. Includes curved-backed knives and some bone tools and ornaments, possibly reflecting Neanderthal acculturation from contact with H. sapiens.

Specific Tool Types

ToolMaterialFunctionTradition
Levallois pointstoneSpear point, cutting toolMousterian
Side scraperstoneHide and wood processingMousterian

Diet & Food Sources

Top-level carnivore; isotopic evidence shows heavy meat diet; also consumed plants, mushrooms, and medicinal herbs

Food SourceTypeEvidenceConfidence
Large game meatMeat huntedIsotopicStrong
Megalafauna marrow (elephant)MarrowIsotopicStrong

Fire Use

100 KYA — Multiple European sites Confirmed

Widespread evidence of habitual fire use for cooking by Neanderthals

70 KYA — Roc de Marsal, France Confirmed

Neanderthal hearths in southwest France.

60 KYA — Kebara Cave, Israel Confirmed

Well-preserved hearths with layered ash deposits

40 KYA — Peştera cu Oase, Romania Probable

Late Neanderthal contexts with fire-maintained occupations.

Shelters & Habitation

Bruniquel Cave structures

176 KYA — Constructed shelter

Neanderthal-constructed ring structures made from approximately 400 broken stalagmites arranged in two rings deep inside a cave. Evidence of fire use on the structures.

Evidence: Two rings of broken stalagmites with fire traces, 336m from cave entrance

Lazaret Cave

130 KYA — Cave

A possible internal tent-like structure within a larger cave, with evidence of hide walls supported against the cave wall.

Evidence: Post-holes along cave wall, concentrated occupation debris within defined area

Roc de Marsal

70 KYA — Rock shelter

Neanderthal occupation with fire.

Evidence: Hearth features

Mezmaiskaya Cave

70 KYA — Cave

Neanderthal infant burial and occupation.

Evidence: Stratified MP layers

Kebara Cave

60 KYA — Cave

Neanderthal hearths and well-preserved occupation.

Evidence: Layered hearths; hyoid burial

Abri du Marais

50 KYA — Rock shelter

MP/UP transition occupations (illustrative).

Evidence: Sheltered overhang

Social Behavior

Shanidar 1 injury survival Confirmed

Healthcare — Shanidar Cave, Iraq

Shanidar 1 Neanderthal survived a crushing blow to the left eye (blinding that eye), a withered right arm (likely amputated), and a severe leg injury. Survival required extended care.

Evidence: Healed fractures and deformities on multiple body regions; individual lived to old age

La Chapelle-aux-Saints care Confirmed

Healthcare — La Chapelle-aux-Saints, France

The "Old Man" of La Chapelle-aux-Saints suffered severe osteoarthritis and tooth loss but survived to old age, implying group support.

Evidence: Severe degenerative disease; survived to estimated 40+ years (old for Neanderthals)

Neanderthal deliberate burials Confirmed

Burial — Multiple sites

Multiple Neanderthal sites show evidence of deliberate burial including flexed body positions, grave goods, and possible flower offerings.

Evidence: Flexed positions in pit-like features; La Ferrassie, Shanidar, Kebara, Teshik-Tash

Neanderthal pyrotechnology Confirmed

Fire use — Europe

Habitual fire in many sites.

Evidence: Hearths across MP

Levallois prepared-core provisioning Probable

Tool sharing — Western Europe

Prepared cores transported across landscapes (Neanderthal sites).

Evidence: Core preparation and flake predetermination

Art & Symbolic Behavior

Neanderthal shell ornaments Confirmed

115 KYA — Cueva de los Aviones, Spain

Perforated and pigment-stained marine shells used by Neanderthals as personal ornaments, predating contact with H. sapiens.

La Ferrassie burials Confirmed

70 KYA — La Ferrassie, France

Multiple Neanderthal burials at La Ferrassie.

Possible Neanderthal cave art Disputed

65 KYA — La Pasiega, Maltravieso, Ardales (Spain)

U-Th dated cave paintings in Spanish caves predating arrival of H. sapiens in Europe (~65 KYA).

Neanderthal manganese processing Probable

60 KYA — Europe

Use of black pigments in some MP sites.

Mousterian abstract marks Disputed

50 KYA — Europe

Engraved objects from some MP sites (debated).

Social Organization

Group Size10–30 individuals
MethodSite area
StructureFission fusion
Sexual Dimorphism1.10x (male/female body mass)
Task DifferentiationSome evidence but less than H. sapiens (Kuhn & Stiner 2006)
TeachingStandardized Mousterian technology implies teaching

Relatively small group sizes based on site areas and genetic evidence of low effective population. Kuhn and Stiner argued for less task differentiation than H. sapiens.

Genetics & Ancient DNA

Genome Coverage99.90%
DNA SourceVindija Cave (Croatia), Altai Mountains (Russia)
Sequencing Year2010
mtDNA AvailableYes
Nuclear DNAYes
Admixture with H. sapiens2.5%
Divergence Date (fossil calibrated)550 KYA
Divergence Date (molecular clock) 0.70 MYA (95% CI: 0.55–0.77; 15 studies)
Key Genes/VariantsFOXP2 gene (human variant), MC1R (red hair variant), TAS2R38 (bitter taste), BNC2 (skin pigmentation), several immune-related HLA variants

First archaic human genome sequenced (Green et al. 2010). High-coverage genome from Altai (Prufer et al. 2014). 1-4% Neanderthal DNA in modern non-Africans.

Molecular clock data from TimeTree 5 (Kumar et al. 2022).

Phylogenetic Relationships

Related SpeciesRelationshipConfidenceNotes
Homo heidelbergensis Proposed ancestor Strong European H. heidelbergensis populations gave rise to Neanderthals
Homo denisova Sister taxon Strong Genomic evidence shows Neanderthals and Denisovans diverged ~400 KYA from a common ancestor
Homo sapiens Interbreeding partner Strong Genomic evidence confirms interbreeding ~50-60 KYA; 1-4% Neanderthal DNA in non-Africans
Homo denisova Interbreeding partner Strong Confirmed by "Denny" — a first-generation hybrid found at Denisova Cave
Homo sp. Nesher Ramla Possible ancestor Moderate Proposed by Hershkovitz et al. as a source population for Neanderthals

Archaeological Evidence

Tools — 176 KYA — Bruniquel Cave, France Confirmed

Neanderthal-constructed ring structures made from broken stalagmites deep inside a cave.

Tools — 125 KYA — Jebel Faya, UAE Probable

Middle Palaeolithic assemblages in Arabia.

Art — 115 KYA — Cueva de los Aviones, Spain Confirmed

Perforated and pigment-stained marine shells used as personal ornaments by Neanderthals.

Healthcare — 50 KYA — Shanidar Cave, Iraq Confirmed

Shanidar 1 survived a crushing blow to the left eye, a withered right arm, and a leg fracture. Survival of these injuries required extended care from others.

Burial — 50 KYA — Shanidar Cave, Iraq Probable

Several Neanderthal skeletons at Shanidar appear to have been deliberately buried, with some associated pollen suggesting possible flower offerings (debated).

Fossils — 49 KYA — El Sidrón Cave, Spain Confirmed

Neanderthal kin group genetics from dental remains.

Fossils — 42 KYA — Vindija Cave, Croatia Confirmed

Neanderthal remains yielding aDNA for genome sequencing.

Art — 40 KYA — Grotte du Renne, France Disputed

Châtelperronian ornaments and bone tools (association debated).

Tools — 40 KYA — Umm el Tlel, Syria Confirmed

Levallois points and MP hunting toolkit.

Key Specimens

SpecimenNameSiteYearAge (MYA)CompletenessSignificance
Shanidar 4 Flower burial Shanidar Cave 1960 0.07 55.00% Famous flower burial evidence (debated)
La Ferrassie 1 1909 0.07 90.00% Classic western European Neanderthal burial
Shanidar 1 Old Man of Shanidar Shanidar Cave 1957 0.07 70.00% Neanderthal pathology and care evidence
Kebara 2 Kebara hyoid 1983 0.06 60.00% Hyoid bone informs speech anatomy debates
La Chapelle-aux-Saints 1 Old Man of La Chapelle 1908 0.05 85.00% Archetypal Neanderthal; severe pathology

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Life History

Estimated Lifespan~45 years
Age at Maturity~15.0 years
Weaning Age~3.0 years
Interbirth Interval~3.5 years
Gestation~9.0 months
Dental DevelopmentSlightly faster than H. sapiens in some histology studies
Brain GrowthLarge Neanderthal brain; high maternal energetic costs
Growth ComparisonSimilar overall pace to modern humans (debated)
ConfidenceEstimated from fossils

Pathology & Healthcare Evidence

Healed fracture — Shanidar 1 Evidence of care

Shanidar Cave — 50 KYA

Affected: Right humerus/clavicle region

Healed crushing fracture with atrophy-related changes; right arm likely impaired.

Care inference: Long-term survival implies provisioning/protection by group.

Survival: Many years post-injury

Arthritis — Shanidar 1 Evidence of care

Shanidar Cave — 50 KYA

Affected: Lower limb joints

Degenerative joint disease; reduced mobility in later life.

Care inference: Survival with limited mobility suggests within-group support.

Survival: Years

Traumatic brain injury — Shanidar 1 Evidence of care

Shanidar Cave — 50 KYA

Affected: Left orbit/face

Blunt trauma to left side of face; healed with vision loss inferred.

Care inference: Survival after severe head trauma.

Survival: Long-term survival

Arthritis — La Chapelle-aux-Saints 1 Evidence of care

La Chapelle-aux-Saints — 50 KYA

Affected: Spine/hip

Severe DJD; osteoarthritis widespread.

Care inference: Old age survival with severe pathology implies social buffering.

Survival: Years

Degenerative — La Ferrassie 1 Evidence of care

La Ferrassie — 50 KYA

Affected: Skeleton

Widespread degenerative changes; advanced age for Neanderthal.

Care inference: Burial context; survival to old age.

Survival: Years

Unhealed fracture — Shanidar 4

Shanidar Cave — 50 KYA

Affected: Rib

Rib lesions interpreted by some as trauma (debated).

Survival: Perimortem vs postmortem debated

Comparative Anatomy

Encephalization Quotient5.10
Intermembral Index78.0
LocomotionStriding biped
Foramen MagnumAnterior
Precision GripModern
Pelvic ShapeBowl shaped
RobusticityRobust
Big ToeFully adducted
ThoraxBarrel shaped

Cold-adapted robust postcrania; large brains (sample ~1200–1750 cc).

Compare anatomy across species →

DNA Data

Mitochondrial / Genomic Sequences

AccessionTypeLength (bp)Organism
KY751400 Complete mitogenome 16,572 Homo neanderthalensis (Vindija 87)
FN673705 Complete mitogenome 16,570 Homo neanderthalensis (El Sidron)
KC879692 Complete mitogenome 16,567 Homo sp. Altai Neanderthal
NC_011137 Complete mitogenome 16,565 Homo sapiens neanderthalensis (Vindija)
FM865411 Complete mitogenome 16,565 Homo neanderthalensis (Mezmaiskaya)
KX198087 Complete mitogenome 16,565 Homo neanderthalensis (Hohlenstein-Stadel)

Ancient DNA Samples

SampleSiteDate (BP)CoverageHaplogroup
Mezmaiskaya1 Mezmaiskaya Cave 70,000 0.50x
Shanidar_Z Shanidar Cave 70,000 0.01x
AltaiNeandertal Denisova Cave 52,000 52.00x
ElSidron1253 El Sidron Cave 49,000 0.30x
Vindija33.19 Vindija Cave 44,000 30.00x
LesCottes_Z4-1514 Les Cottes 42,000 2.70x

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Isotope Analyses

SystemValueMaterialSiteDate (MYA)Interpretation
delta N15 10.20 Bone collagen Qesem Cave 0.400 Qesem Cave: late Lower Palaeolithic meat-dominated protein signal (site studies).
delta N15 10.80 Bone collagen Schoningen 0.300 European Middle Palaeolithic carnivory signal (site-specific).
delta N15 10.50 Bone collagen 0.050 Neanderthals: high δ15N consistent with high trophic level meat protein.
delta N15 9.10 Bone collagen 0.050 Lower δ15N end of Neanderthal range (ecology/age effects).
delta N15 11.20 Bone collagen Shanidar Cave 0.045 Shanidar: elevated δ15N in some individuals (freshwater caveat in some regions).

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Dating Evidence

MethodDate (MYA)UncertaintyMaterialSite / Specimen
Ar Ar 1.2000 ± 0.0500 Atapuerca section tephras Atapuerca
U series 0.4300 ± 0.0200 Speleothems / flowstone (Sima) Atapuerca / Cranium 5
U series 0.3000 ± 0.0300 Speleothem / peat (context) Schoningen
Radiocarbon C14 0.0500 ± 0.0050 Charcoal (Neanderthal layers) Shanidar Cave / Shanidar 1
Radiocarbon C14 0.0500 ± 0.0100 Charcoal / bone (molecular age) Denisova Cave
Radiocarbon C14 0.0400 ± 0.0050 Charcoal (Aurignacian contexts) Hohle Fels

Fossil Occurrences

The Paleobiology Database records 6 fossil occurrence(s) attributed to Homo neanderthalensis. View on map →

Identified AsLocationFormationAge (MYA)
Homo neanderthalensis GR 0.77 – 0.01
Homo cf. neanderthalensis FR 0.77 – 0.13
Homo neanderthalensis GR 0.13 – 0.01
Homo neanderthalensis GR 0.13 – 0.01
Homo neanderthalensis PT 0.13 – 0.01
Homo neanderthalensis FR 0.13 – 0.01

Data from the Paleobiology Database (CC-BY).

Scientific References

  1. Hoffmann DL, Standish CD, Garcia-Diez M, et al. (2018). "U-Th dating of carbonate crusts reveals Neandertal origin of Iberian cave art". Science 359:912-915. DOI:10.1126/science.aap7778 (395 citations)
  2. Neubauer S, Hublin JJ, Gunz P (2018). "The evolution of modern human brain shape". Science Advances 4:eaao5961. DOI:10.1126/sciadv.aao5961 (283 citations)
  3. Spikins P, Needham A, Tilley L, Hitchens G (2018). "Calculated or caring? Neanderthal healthcare in social context". World Archaeology 50:384-403. DOI:10.1080/00438243.2018.1433060 (41 citations)
  4. Zilhao J, Angelucci DE, Badal-Garcia E, et al. (2018). "Symbolic use of marine shells and mineral pigments by Iberian Neandertals 115,000 years ago". Science Advances 4:eaar5255. DOI:10.1126/sciadv.aar5255 (128 citations)
  5. Prufer K, Racimo F, Patterson N, et al. (2014). "The complete genome sequence of a Neanderthal from the Altai Mountains". Nature 505:43-49. DOI:10.1038/nature12886 (2,023 citations)
  6. Sankararaman S, Mallick S, Dannemann M, et al. (2014). "The genomic landscape of Neanderthal ancestry in present-day humans". Nature 507:354-357. DOI:10.1038/nature12961 (954 citations)
  7. Green RE, Krause J, Briggs AW, et al. (2010). "A draft sequence of the Neandertal genome". Science 328:710-722. DOI:10.1126/science.1188021 (3,599 citations)
  8. Arensburg B, Tillier AM, Vandermeersch B, et al. (1989). "A Middle Palaeolithic human hyoid bone". Nature 338:758-760