Overview

The first hominin species ever recognized, described by Raymond Dart in 1925 from the Taung Child skull. Known from numerous specimens from South African cave sites including Sterkfontein ("Mrs. Ples" STS 5) and Makapansgat. Had a larger brain relative to body size than Au. afarensis. Evidence of a more varied diet and potential early stone tool use, though this is debated.

Key Fossils

Taung 1 (Taung Child), STS 5 ("Mrs. Ples"), STS 71, StW 573 ("Little Foot")

Brain Anatomy

Endocast-Based Estimates

Brain region sizes are estimated from endocasts (internal skull casts), not direct brain observation. These are approximations with inherent uncertainty.
Frontal
149 cc
Parietal
99 cc
Temporal
90 cc
Occipital
77 cc
Cerebellar
36 cc
RegionFunctional Implicationsvs. Modern Human
Frontal Modest prefrontal expansion compared to apes; basic planning and social cognition Smaller proportionally; less prefrontal development
Parietal Expanded compared to apes; spatial processing and tool-related cognition Smaller; less parietal integration
Temporal Similar to ape proportions; auditory and memory processing Less developed temporal association areas
Occipital Relatively large occipital lobe; strong visual processing Larger proportionally than modern humans (more visual emphasis)
Cerebellar Relatively small cerebellum Proportionally smaller

Tools & Technology

Specific Tool Types

ToolMaterialFunctionTradition
Digging stickwoodExtracting underground storage organs (tubers)

Diet & Food Sources

Varied diet; isotopic evidence suggests mix of C3 and C4 foods, possibly including some animal protein

Food SourceTypeEvidenceConfidence
Tubers and roots (C3)TuberIsotopicModerate

Social Organization

Group Size25–45 individuals
MethodDunbar neocortex
StructureMulti male multi female
Sexual Dimorphism1.35x (male/female body mass)

Au. africanus: moderate dimorphism suggests multi-male groups.

Phylogenetic Relationships

Related SpeciesRelationshipConfidenceNotes
Australopithecus afarensis Proposed ancestor Moderate Au. afarensis is a likely ancestor of the South African Au. africanus
Homo habilis Possible ancestor Weak One of several proposed routes to the origin of Homo
Australopithecus prometheus Sister taxon Weak Debated; Clarke argues distinct from A. africanus; critics see them as conspecific

Key Specimens

SpecimenNameSiteYearAge (MYA)CompletenessSignificance
Taung 1 Taung Child Sterkfontein 1924 2.80 55.00% First African hominin fossil
STS 5 Mrs. Ples Sterkfontein 1947 2.50 85.00% Most complete A. africanus cranium
Stw 573 Little Foot Sterkfontein 1994 2.20 90.00% Among most complete early hominin skeletons

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Life History

Estimated Lifespan~32 years
Age at Maturity~11.5 years
Weaning Age~3.5 years
Interbirth Interval~4.3 years
Gestation~8.0 months
Dental DevelopmentTaung dental eruption (historic estimates)
Brain GrowthModerate brain for body
Growth ComparisonIntermediate between chimps and Homo
ConfidenceEstimated from fossils

Pathology & Healthcare Evidence

Nutritional stress — Taung 1

Taung — 2.8 MYA

Affected: Enamel

Enamel hypoplasia lines in Taung hominin (historic interpretations).

Survival: Early childhood

Healed fracture — Stw 573

Sterkfontein — 2.7 MYA

Affected: Elements of StW 573

Some healed postcranial trauma reported in Little Foot assessments (site-specific claims).

Survival: Healed

Comparative Anatomy

Encephalization Quotient2.65
Intermembral Index92.0
LocomotionObligate biped
Foramen MagnumAnterior
Precision GripRudimentary
Pelvic ShapeIntermediate
RobusticityModerate
Big ToePartially adducted
ThoraxFunnel shaped

Au. africanus: biped with climbing retention.

Compare anatomy across species →

Isotope Analyses

SystemValueMaterialSiteDate (MYA)Interpretation
delta C13 -6.50 Enamel Sterkfontein 2.800 Sterkfontein Au. africanus: mixed C3/C4 browsing/grazing mosaic.
delta C13 -8.20 Enamel Sterkfontein 2.500 Lower end of Au. africanus δ13C distribution (South African sites).
delta O18 -4.20 Enamel Sterkfontein 2.500 Bioapatite carbonate oxygen (reporting conventions vary; illustrative site-level value).

Explore all isotope and climate data →

Dating Evidence

MethodDate (MYA)UncertaintyMaterialSite / Specimen
Cosmogenic nuclide 2.8000 ± 0.3000 Member 4 breccia (surface exposure models) Sterkfontein
U Pb 2.8000 ± 0.0500 Flowstone (Taung) Middle Awash / Taung 1
U Pb 2.6700 ± 0.1500 Flowstone (Little Foot) Sterkfontein / Stw 573

Fossil Occurrences

The Paleobiology Database records 7 fossil occurrence(s) attributed to Australopithecus africanus. View on map →

Identified AsLocationFormationAge (MYA)
Australopithecus cf. africanus KE Nawata and Nachukui 11.63 – 3.60
Australopithecus cf. africanus KE Lothagam 1 7.25 – 2.58
Australopithecus africanus ET USNO 5.33 – 2.58
Australopithecus africanus ZA Sterkfontein 5.33 – 0.77
Australopithecus africanus ZA Makapansgat 5.33 – 2.58
Australopithecus africanus ZA 5.33 – 2.58
Australopithecus africanus ZA Makapansgat 3.60 – 2.58

Data from the Paleobiology Database (CC-BY).

Scientific References

  1. Dart RA (1925). "Australopithecus africanus: The Man-Ape of South Africa". Nature 115:195-199